
I used a programs anti-aliasing tool on this with inverts from the color wheel. It looks like dark outlines on light colors because of color brilliance. In black and white, even though with the red/green, they have the exact same value/luminance.
To me, there is no solution to blending inverts. I'd go with really smooth splines or lines without anti-aliasing to do this if I had to. Also, if you look closely, the anti-aliasing seems to extrude the circles to the center like buttons. Look closer, and the anti-aliasing is mirrored, so this is just an illusion. Probably due to the brilliance of yellow drawing the eye's focus.
Edit:

If you look at the gradient (cg), the colors go through a colors that look like crap, and which seem darker in the middle.