The last WIP was a huge step up, and also rather unique, imo, since I normally associate large, round, cartoony shapes with that you find on a title screen. It's rare to see someone being able to fit that into the game itself! Very nice!

I think that the frog character does blend a bit too much into the background, though.

I've tried to improve on the ledge crawl.

This animation rubs me the wrong way, unfortunately. It has the same feeling as bad old 16-bit platformers, and I'll tell you why:

1) The action stops dead for a millisecond (when he catches the ledge, everything freezes or he gets stuck to it).
2) There is no elasticity (all of the action happens at roughly the same, close to linear speed).

The easy way I would fix this would be to simply make his feet quickly bounce out and in a pixel during the dead frames when he catches the ledge. I think just that would help to make it so much more alive (also think of all the elasticity the old Super Mario Games had, which not many other platformers were able to mimic).

PS. Love the idea btw (reminds me of Ramrod, somehow, which sadly never was released), and hope to see an OS X version too?

To continue my feedback... I don't know if people will confuse the monkey with the lion since they do look kinda similar, but I'd guess that people might think that the monkey and lion are on the same sides, since they share the same color.

How about replacing the lion with either a tiger (essentially a walking warning sign with the stripes... heck, a bee/wasp would do) or a panther (though people might confuse that with a black cat, since the tiles are all the same size)?

This feels a bit like adventures of lolo now as well, come to think of it. Wasn't that similar to chip's challenge?

I would love to see someone do an edit with white blades on the scythe/axe, just so it might balance the wings a bit.

It's an incredibly impressive piece!