From what i gathered, Udon crew made some keyframe for each sprite, then another studio called Studio XD continued to make the rest. The pictures posted here look similar to those in a comic called "Echo" which was pencilled by a guy called Long Vo. So even though Udon certainly has people that could do this right, it's this Long Vo person who contributes to this weird anatomy, pillowshading and so forth. All those features can also be found in his comic.