I have to admit, I've been feeling a bit reluctant to change it dramatically at this stage, and there are still some things I'm not sure how to fix, but I hope this takes care of the most important things.
OK, first of all, I increased the saturation of the brightest skin shades to make up for the green AA, so now the colors should seem less muddled. Added another skin shade a bit closer to green.
Not sure if the wings need changing since they're meant to be damaged rather than neat and symmetrical.
He does look less scary in a way, but I thought the old one looked scary in a cornered beast sort of way and that maybe something more intelligent would seem more threatening. I guess it's a matter of how you choose to read it?
Indigo, that makes perfect sense, and I seem to have more trouble keeping the more basic stuff in mind when trying more challenging things.
Anyhow, thanks a lot for another helpful edit.
Edit: Nothing major, mostly palette changes and polishing.