I think it's fine to necro your own thread :]
https://i.imgur.com/iXHc8VZ.pngI think this looks nice!
The colours feel a little dull and could use more hue-shifting, maybe something like this?
(I seem to have introduced a bunch of extra colours while recolouring it with blending modes, please don't mind that.)
Try making your vertical/horizontal edge tiles less straight, make them more winding, give them a more interesting silhouette. Right now, in the example, they make the tile grid very obvious because they're so straight. Same with the edges of the corners, to a lesser degree.
There are some single light-coloured pixels in the interior, they look like noise rather than like leaves. I'd get rid of them.
I have a hard time believing you need that many variants of the dark interior tiles xP They all feel samey since they use the same leaf shapes and similar arrangements, so you can probably use fewer of them and achieve the same effect. Or, you can add more variety by having some thicker clumps, and maybe even some tiles that use the next colour up the ramp.
The bottoms of the thick trunks feel a bit off for a couple of reasons:
- the roots slope down too vertically on the sides, making the tree look like it's standing on its roots rather than like we're seeing the parts of the roots that are visible above the ground.
- the roots have very similar sizes and shapes. Try to vary them more, in particular the shapes.