This is comming along nicely. I can tell you are really putting effort into this.
Some crits. You have a few colours which are a bit close, not many but still. It looks you as if you aim for a 32 colour limit and in that case you wanna utilise all your colours to maximum effect.
In some places where you started to aa it is too much, A general rule of thumb i go by is make things smooth but keep em sharp, too much aa gets blurry.
In some of the gadiations you dont you all the colours you could to make it look smooth and thus the dither looks a tad harsh in places.
Also the blue you used to go between the green in some places does not really work for me, it is good as an interchangable hue for one of the darker greens tho.
Lot's of aa and gradiation issues can be fixed by just keeping a neat palette in promotion (and i know you use promotion that's why i say it
)
So doing aa and colour gradiations quick with using [ ] is easier in most cases unless you have superwild hueshifting and lots of intersecting ramps going on, in that case i suggest workramps where you just put new ramps in your palette with the same colours you already have but in different working gradiations.
I also agree with rob about the wrinkles.
Here a lil edit to illustrate some of my points, i also changed the colours a bit and neatened the palette (in case you wanna look at it in pm)
The colouredits are very rough tho and could be way nicer with better tints of the several palettes.
and animation to compare to yours
keep it up