I belive you are focusing too much on the independent feathers, it is more important to make the structure clear
http://www.birding.com/IDwing.asp http://www.birding.com/IDbody.aspthe biggest glaring error is the way you emphasized their borders, they dont really go trough the whole extension of the wing, they only go up to the wrist.
detailing a feather so much is only any use with the primary feathers, and even then I belive it should be done much more subtly, by going up/down a shade if anything.
The wings are also shaded in a borderline pillowshading way, which helps even more to make them look tacky. you shouldnt shade the lit and the shadowed parts the same way, because it flatterns the image, you need to decide on focus points so you can put the detail accordingly.
the bone structure is really weird on his right wing, it looks like it'd need an extra bone to bend the way it is while it does from the shoulder to the wrist.
Regarding the nose/eye in shadow, if you can honestly mistake an eye for a nose, there has to be something wrong with the picture. the lone eyes in the shadow should be mysterious and intriguing from a "what is he thinking" point of view, not a "what the hell is that supposed to be?" point of view.